The Evo-X engine has cylinders that are separate castings from the lower case and the heads, and cast in iron liners for the cylinders. On the V-Rod engine, the aluminum cases extend all the way to the head and the cylinders are steel wet sleeves held in the cases with o-rings, much like those in a big diesel engine.
Another difference between the V-Rod and this engine are the cam drives. If you look carefully at the cylinder heads you can see the front head has it's sparkplug located on the left side of the head casting while the rear cylinder has the spark plug located on the right side of the head casting. This tells us that the cam chain for the front cylinder is on the right side of the block and the cam chain for the rear cylinder is on the left side of the block. This is exactly the same as a Yamaha Virago, Buell Helicon engine, several Rotax watercraft and snowmobile engines and before any of these, the Harley Davidson 1100 OHC prototype that never made production. V-Rod engines place both cam drives on the left side of the engine and the spark plugs are centrally located in the combustion chamber.
If you look carefully, it appears to me the front and rear heads use the same casting but they are simply oriented 180 degrees apart on their respective cylinders. This would make a lot of sense from a cost savings standpoint. V-Rods use dual down draft throttle bodies and velocity stacks with an injector in each intake. From what I can read about the Street, there is a single TB and one injector for both cylinders, a huge kludge in my opinion.
You can also see a coil on the left side of the bike where Harley usually places the horn. V-Rods have coils built into the spark plug cap. Coolant hose routing is very different from that of a V-Rod so I have to suspect a different coolant pump location.
And the original VR-1000 race bike engine, from which Porsche Engineering developed a workable and reliable steet engine for the V-Rod.